caa a22 4500 388017066 CHVBK 20180307124914.0 cr unu---uuuuu 161130e199809 xx s 000 0 eng 10.1017/S0022336000027189 doi S0022336000027189 pii (NATIONALLICENCE)cambridge-10.1017/S0022336000027189 The classification of athyridid brachiopods [Elektronische Daten] [Fernando Alvarez, Rong Jia-Yu, A. J. Boucot] Phylogenetic relationships among higher taxa of articulate brachiopods that developed a spiralium as mineralized support for the lophophore, and the resulting classifications, are not universally agreed upon due to the complex pattern of character evolution within the phylum. Our results, using phylogenetic methodology in conjunction with traditional methods, showed a more complete and easily testable picture of this evolution than either method used alone. A detailed working hypothesis of athyridide phylogeny and the supporting evidence on character distribution is given, and a new classification of the impunctate and punctate athyrids sensu lato is presented with diagnoses to subfamily. Agreement between the stratigraphical first appearance of athyridid subfamilies and their cladistic rank was quite good (except for the koninckinoids), suggesting that both outgroup and traditional paleontological methods indicate a similar direction of character polarity in the evolution of the group. The classification is proposed based on a mix of external and internal characters which persisted during unbroken lineages. New taxa are the families Parazygidae and Triathyrididae; the subfamilies Whitfieldellinae, Triathyridinae, Septathyridinae, Cleiothyridininae, Ochotathyridinae and Plectospirinae, and the genera Brimethyris, Carteridina, Tetraloba and Cooperispira. We studied the great morphological diversity among the athyridids. Their complete stratigraphical ranges help in building phyletic lineages. Copyright © The Paleontological Society Alvarez Fernando Departamento de Geología, Universidad de Oviedo, Oviedo 33005, Spain aut Jia-Yu Rong Nanjing Institute of Geology and Palaeontology, Academia Sinica, Chi-Ming-Ssu, Nanjing 210008, People's Republic of China aut Boucot A. J. Department of Zoology, Oregon State University, Corvallis, Oregon 97331 aut Journal of Paleontology Cambridge University Press 72/5(1998-09), 827-855 0022-3360 72:5<827 1998 72 JPA https://doi.org/10.1017/S0022336000027189 text/html Onlinezugriff via DOI 1 research-article jats NATIONALLICENCE 856 40 https://doi.org/10.1017/S0022336000027189 text/html Onlinezugriff via DOI NATIONALLICENCE 700 1- Alvarez Fernando Departamento de Geología, Universidad de Oviedo, Oviedo 33005, Spain aut NATIONALLICENCE 700 1- Jia-Yu Rong Nanjing Institute of Geology and Palaeontology, Academia Sinica, Chi-Ming-Ssu, Nanjing 210008, People's Republic of China aut NATIONALLICENCE 700 1- Boucot A. J. Department of Zoology, Oregon State University, Corvallis, Oregon 97331 aut NATIONALLICENCE 773 0- Journal of Paleontology Cambridge University Press 72/5(1998-09), 827-855 0022-3360 72:5<827 1998 72 JPA CC0 http://creativecommons.org/publicdomain/zero/1.0 nationallicence BK010053 XK010053 XK010000 NATIONALLICENCE NATIONALLICENCE NL-cambridge