caa a22 4500 388017384 CHVBK 20180307124914.0 cr unu---uuuuu 161130e199801 xx s 000 0 eng 10.1017/S0022336000023982 doi S0022336000023982 pii (NATIONALLICENCE)cambridge-10.1017/S0022336000023982 Color banding in the Triassic terebratulid brachiopod Coenothyris from the Muschelkalk of Central Europe [Elektronische Daten] [Hans Hagdorn, Michael R. Sandy] The Triassic terebratulid brachiopod Coenothyris frequently displays preserved color patterns; such patterns have commonly been recorded from Paleozoic terebratulid brachiopods. Despite the frequency with which color patterns are preserved in Coenothyris, there has been no recent investigation of the cause and significance of this phenomenon. Shell material is well-preserved; energy-dispersive spectroscopy and microprobe analysis has been unable to detect compositional differences between colored and noncolored shell. This supports the organic origin of the color patterns as suggested for Devonian terebratulids by Richter (1919); color patterns originate from organic pigment in the primary shell layer. Three subtypes of radial color banding are identified: subtype A with a relatively large number (up to 80) of delicate color bands on each valve; subtype B with fewer and generally wider color bands (less than 20) on each valve of adult specimens; subtype C with faint to fairly wide but very short color bands along the anterior margin (ranging from a few to more than 50 in number). Serial sections prepared from subtypes A and B confirm their congeneric status. As shell form (length/width/thickness ratio) and maximum size varies, color pattern types differ in various stratigraphic horizons and also in isochronous populations from different geographic localities, indicating different facies. However, the variation in color patterns is not due to systematic differences at the species or subspecies level but rather reflects a tendency among Coenothyris vulgaris to respond to different environmental parameters. This variation in color patterns is ecophenotypic. Copyright © The Paleontological Society Hagdorn Hans Muschelkalkmuseum, Schloßstraße 11, 74653 Ingelfingen, Germany, <Encrinus@t-online.de> aut Sandy Michael R. Department of Geology, University of Dayton, Dayton, Ohio 45469-2364, U.S.A., <sandy@neelix.udayton.edu> aut Journal of Paleontology Cambridge University Press 72/1(1998-01), 11-28 0022-3360 72:1<11 1998 72 JPA https://doi.org/10.1017/S0022336000023982 text/html Onlinezugriff via DOI 1 research-article jats NATIONALLICENCE 856 40 https://doi.org/10.1017/S0022336000023982 text/html Onlinezugriff via DOI NATIONALLICENCE 700 1- Hagdorn Hans Muschelkalkmuseum, Schloßstraße 11, 74653 Ingelfingen, Germany, <Encrinus@t-online.de> aut NATIONALLICENCE 700 1- Sandy Michael R. Department of Geology, University of Dayton, Dayton, Ohio 45469-2364, U.S.A., <sandy@neelix.udayton.edu> aut NATIONALLICENCE 773 0- Journal of Paleontology Cambridge University Press 72/1(1998-01), 11-28 0022-3360 72:1<11 1998 72 JPA CC0 http://creativecommons.org/publicdomain/zero/1.0 nationallicence BK010053 XK010053 XK010000 NATIONALLICENCE NATIONALLICENCE NL-cambridge